Photosynthesis pigments and antioxidants were measured in field-collected plants from all treatments as well as in lab-cultured plants. Zeaxanthin and chlorophyll a:b levels increased with UV-filtering, while neoxanthin and chlorophyll b decreased (Fig. 4). Un-manipulated site reference samples also had significantly more zeaxanthin, lutein, β-carotene, and a higher chlorophyll a:b, as well as lower violaxanthin, neoxanthin, chlorophyll a, and chlorophyll b than UV-transmitted samples. There was no significant difference in violaxanthin, antheraxanthin, lutein, β-carotene, or chlorophyll a between UV-filtered samples and UV-transmitted samples, though the VAZ pool was larger in UV-filtered plants than in UV-transmitted plants (P = 0.048).