Acknowledgements
This work was supported by the University of California, Berkeley, Department of Integrative Biology Graduate Research Fund to JTBE; the UC Natural Reserve System Mildred E. Mathias Research Grant to JTBE; the American Bryological and Lichenological Society Anderson & Crum Field Research Award to JTBE; German Research Foundation (DFG) project number 427925948 to OD; and National Science Foundation Dimensions of Biodiversity awards (DEB-1638956 and DEB 1638943) to BDM and LRS, respectively. KKN is an investigator of the Howard Hughes Medical Institute. JTBE was also supported by the University of California Berkeley Fellowship, the UC Berkeley Pinto-Fialon Fellowship, and the National Science Foundation Dimensions of Biodiversity award (DEB-1638956).