Author contributions
SZ, AQ, TW, LF, JJ, GD conceived research. SZ, AQ, OKB, SM, ASH, GD, TW, TL performed research. SZ, AQ, OKB, SM, ASH, YS, TC, JJ, TW, GD analyzed data. SZ and GD wrote the manuscript. All the authors edited and accepted the final version of the manuscript.