References
Abdelkrim, J., Aznar-Cormando, L, Buge, B., Fedosov, A., Kantor, Y.,
Zaharias, P, and Puillandre, N. (2018) Delimiting species of marine
gastropods (Turridae, Conoidea) using RAD sequencing in an integrative
taxonomy framework. Molecular Ecology, 27, 4591-4611.
Ahmed, M. Z., Araujo-Jnr, E. V., Welch, J. J. and Kawahara, A. Y. (2015)
Wolbachia in butterflies and moths: geographic structure in infection
frequency. Frontiers in Zoology, 12, 1-9.
Andersson, L. (1990) The driving force: species concepts and ecology.
Taxon, 39(3), 375-382.
Andrews, K. R., Good, J. M., Miller, M. R., Luikart, G., and Hohenlohe,
P. 2016. Harnessing the power of RADseq for ecological and evolutionary
genetics. Nature Reviews Genetics , 17 , 81-92.
Austin, M. P. (1985) Continuum concept, ordination methods, and niche
theory. Annual review of ecology and systematics, 16(1), 39-61.
Baird, N. A., Etter, P. D., Atwood, T. S., Currey, M. C., Shiver, A. L.,
Lewis, Z. A., Selker, E. U., Cresko, W. A., and Johnson, E. A. (2008)
Rapid SNP discovery and genetic mapping using sequenced RAD markers.
PLoS ONE, 3, e3376.
Balkenhol, N., Cushman, S., Storfer, A., Waits, L. (2016) Landscape
Genetics: Concepts, Methods, Applications. Wiley, West Sussex, UK.
Bandelt, H., Forster, P., and Röhl, A. (1999) Median-joining networks
for inferring intraspecific phylogenies. Molecular Biology and
Evolution, 16, 37-48.
Bird, C. D., Hilchie, G. J., Kondla, N. G., Pike, E. M., and Sperling,
F. A. H. 1995. Alberta Butterflies. The Provincial Museum of Alberta,
Edmonton, Alberta, Canada.
Bond, J. E., & Stockman, A. K. (2008) An integrative method for
delimiting cohesion species: finding the population-species interface in
a group of Californian trapdoor spiders with extreme genetic divergence
and geographic structuring. Systematic Biology, 57(4), 628-646.
Bouckaert, R. (2010) DensiTree: making sense of sets of phylogenetic
trees. Bioinformatics, 26, 1372-1373.
Bouckaert, R., Heled, J., Kühnert, D., Vaughan, T., Wu, C., Xie, D.,
Suchard, M. A., Rambaut, A., and Drummond, A. J. (2014) BEAST 2: a
software platform for Bayesian evolutionary analysis. PLoS Computational
Biology, 10, e1003537.
Brock, J. P. and Kaufman, K. (2003) Kaufman Field Guide to Butterflies
of North America, New York: Houghton Mifflin Co.
Brooks ME, Kristensen K, van Benthem KJ, Magnusson A, Berg CW, Nielsen
A, Skaug HJ, Maechler M, Bolker BM (2017). “glmmTMB Balances Speed and
Flexibility Among Packages for Zero-inflated Generalized Linear Mixed
Modeling.” The R Journal, 9(2), 378–400.
Bryant, D., Bouckaert, R., Felsenstein, J., Rosenberg, N. A., and
Choudhury, A. R. (2012) Inferring species trees directly from biallelic
genetic markers: bypassing gene trees in a full coalescent analysis.
Molecular Biology and Evolution, 29, 1917-1932.
[dataset] Campbell, E. O., MacDonald, Z. G., Gage, E. V., Gage, R.
V., and Sperling, F. A. H. (2020). Data from: Reconciling genomic and
ecological species delimitation using a confusing group of butterflies.
NCBI accessions NNNN-NNNN.
Campbell, E. O., Davis, C. S., Dupuis, J. R., Muirhead, K., and
Sperling, F. A. H. (2017) Cross-platform compatibility of de
novo-aligned SNPs in a nonmodel butterfly genus. Molecular Ecology
Resources, 6, e84-e93.
Campbell, E. O., Gage, E., V., Gage, R. V., and Sperling, F. A. H.
(2019) SNP-based species phylogeny of greater fritillary butterflies
(Lepidoptera: Nymphalidae: Speyeria) demonstrates widespread
mito-nuclear discordance. Systematic Entomology, DOI:10.1111/syent.12393
Carstens, B. C., Pelletier, T. A., Reid, N. M., and Satler, J. D. (2013)
How to fail at species delimitation. Molecular Ecology, 22, 4369-4383.
Catchen, J. M., Amores, A., Hohenlohe, P., Cresko, W., and Postlethwait,
J. H. (2011) Stacks: building and genotyping loci de novo from
short-read sequences. G3, 1, 171-182.
Chandra, A.K., Raghavan, P., Ruzzo, W.L., Smolensy, R. & Tiwari, P.
1996. The electrical resistance of a graph captures its commute and
cover times. Computational Complexity, 6(4), 312-340.
Chen, C., Durand, E., Forbes, F., and François, O. (2007) Bayesian
clustering algorithms ascertaining spatial population structure: a new
computer program and a comparison study. Molecular Ecology Notes, 7,
747-756.
Coates, D. J., Byrne, M., and Moritz, C. (2018) Genetic diversity and
conservation units: dealing with the species-population continuum in the
age of genomics. Frontiers in Ecology and Evolution, 6, 165.
Coyne, J.A. & Orr, H.A. (2004). Speciation. Sinauer Associates,
Sunderland, MA.
Crispo, E., Bentzen, P., Reznick, D.N., Kinnison, M.T. & Hendry, A.P.
(2006) The relative influence of natural selection and geography on gene
flow in guppies. . Molecular Ecology, 15, 49-62.
Danecek, P., Auton, A., Abecasis, G., Albers, C. A., Banks, E.,
DePristo, M. A., … 1000 Genomes Analysis Group. (2011) The
variant call format and VCFtools. Bioinformatics, 27, 2156-2158.
de Moya, R. S., Savage, W. K., tenney, C.., Bao, X., Wahlberg, N., and
Hill, R. I. 2017. Interrelationships and diversification ofArgynnis Fabricius and Speyeria Scudder butteflies.Systematic Entomology , 42 , 635-649.
de Queiroz, K. (1998) The general lineage concept of species, species
criteria, and the process of speciation. IN Howard, D. J., Berlocher, S.
H. (Eds.) Endless forms: species and speciation. Oxford University
Press, New York.
de Queiroz, K. (2007) Species Concepts and Species Delimitation.
Systematic Biology, 56, 879-886.
dos Passos, C. F., and Grey, L. P. (1947) Systematic catalogue of
Speyeria (Lepidoptera, Nymphalidae) with designations of types and
fixations of type localities. American Museum Novitates, 1370, 1-30.
Drummond, A. J., and Rambaut, A. (2007) BEAST: Bayesian evolutionary
analysis by sampling trees. BMC Evolutionary Biology, 7: 214.
Dunford, J. C. (2009) Taxonomic overview of the greater fritillary genus
Speyeria Scudder and the atlantis-hesperis species complexes, with
species accounts, type images, and relevant literature (Lepidoptera:
Nymphalidae). Insecta Mundi, 90, 1-74.
Durand, E., Jay, F., Gaggiotti, O. E., and François, O. (2009) Spatial
inference of admixture proportions and secondary contact zones.
Molecular Biology and Evolution, 14, 2611-2620.
Elith*, J., H. Graham*, C., P. Anderson, R., Dudík, M., Ferrier, S.,
Guisan, A., … & Li, J. (2006). Novel methods improve prediction of
species’ distributions from occurrence data. Ecography, 29(2), 129-151.
Elith, J., Phillips, S. J., Hastie, T., Dudík, M., Chee, Y. E., &
Yates, C. J. (2011). A statistical explanation of MaxEnt for ecologists.
Diversity and Distributions, 17(1), 43-57.
Escudero, M., Eaton, D. A. R., Hahn, M., Hipp, A. L. (2014)
Genotyping-by-sequencing as a tool to infer phylogeny and ancestral
hybridization: a case study in Carex (Cyperaceae). Molecular
Phylogenetics and Evolution, 79, 359-367.
Evanno, G., Regnaut, S., and Goudet, J. (2005) Detecting the number of
clusters of individuals using the software Structure: A simulation
study. Molecular Ecology, 14, 2611-2620.
Freudenstein, J. V., Broe, M. B., Folk, R. A., & Sinn, B. T. (2016).
Biodiversity and the species concept—Lineages are not enough.
Systematic Biology, syw098.
Graham, C. H., Ron, S. R., Santos, J. C., Schneider, C. J., & Moritz,
C. (2004) Integrating phylogenetics and environmental niche models to
explore speciation mechanisms in dendrobatid frogs. Evolution, 58(8),
1781-1793.
Grey, L. P. (1951) The subspeciation of Speyeria atlantis. The
Lepidopterist’s News, 5, 31-35.
Grummer, J. A., Bryson, R. W. Jr., and Reeder, T. W. (2014) Species
delimitation using Bayes factors: simulations and applications to the
Sceloporus scalarsis species group (Squamata: Phrynosomatidae).
Systematic Biology, 63, 119-133.
Guppy, C. S., and Shepard, J. H. (2001) Butterflies of British Columbia.
UBC Press; Vancouver, British Columbia. 414p.
Guisan, A., & Zimmermann, N. E. (2000). Predictive habitat distribution
models in ecology. Ecological modelling, 135(2-3), 147-186.
Hamm, C. A., Handley, C. A., Pike, A., Forister, M. L., Fordyce, J. A.,
and Nice, C. C. (2014) Wolbachia infection and Lepidoptera of
conservation concern. Journal of Insect Science, 14:6.
Hammond, P. C., McCorkle, D., and Bergman, W. (2013) Hybridization
studies of genomic compatibility and phenotypic expression in the
greater fritillary butterflies (Nymphalidae: Argynnini). Journal of the
Lepidopterists’ Society, 67, 263-273.
Hijmans, R.J, Phillips, S., Leathwick, J. and Elith, J. (2011). Package
‘dismo’. Available online at:
http://cran.r-project.org/web/packages/dismo/index.html.
Hijmans, R.J., & van Etten, J. (2012). raster: Geographic analysis and
modeling with raster data. R package version 2.0-12.
http://CRAN.R-project.org/package=raster.
Hinojosa, J. C., Koubínová, D., Szenteczki, M. A., Pitteloud, C., Dincă,
V., Alvarez, N., and Vila, R. (2019) A mirage of cryptic species:
genomics uncover striking mitonuclear discordance in the butterfly
Thymelicus sylvestris. Molecular Ecology. DOI:
https://doi.org/10.1111/mec.15153
Hoang, D. T., Chernomor, O., von Haeseler, A., Ming, B. Q., and Vinh, L.
S. (2018) UFBoot2: Improving the ultrafast bootstrap approximation.
Molecular Biology and Evolution, 35, 518-522.
Hohenlohe, P. A., Day, M. D., Amish, S. J., Miller, M. R., Kamps-Hughes,
N., Boyer, M. C., Muhlfeld, C. C., Allendorf, F. W., Johnson, E. A., and
Luikart, G. (2013) Genomic patterns of introgression in rainbow and
westslope cutthroat trout illuminated by overlapping paired-end RAD
sequencing. Molecular Ecology, 22, 3002-3013.
Hundsdoerfer, A., K., Lee, K. M., Kitching, I. J., and Mutanen, M.
(2019) Genome-wide SNP data reveal an overestimation of species
diversity in a group of hawkmoths. Genome Biology and Evolution, 11,
2136-2150.
Hurst, G. D. D. and Jiggins, F. M. (2005) Problems with mitochondrial
DNA as a marker in population, phylogeographic and phylogenetic studies:
the effects of inherited symbionts. Proceedings of the Royal Society B,
272, 1525-1534.
IPBES (2019) Global assessment report on biodiversity and ecosystem
services of the Intergovernmental Science-Policy Platform on
Biodiversity and Ecosystem Services. E. S. Brondizio, J. Settele, S.
Díaz, and H. T. Ngo (editors). IPBES secretariat, Bonn, Germany.
Jezkova, T., Jaeger, J. R., Marshall, Z. L., & Riddle, B. R. (2009)
Pleistocene impacts on the phylogeography of the desert pocket mouse
(Chaetodipus penicillatus ). Journal of Mammalogy, 90(2), 306-320.
Kodandaramaiah, U., Simonsen, T. J., Bromilow, S., Wahlberg, N., and
Sperling, F. A. H. (2013) Deceptive single-locus taxonomy and
phylogeography: Wolbachia-associated divergence in mitochondrial DNA is
not reflected in morphology and nuclear markers in a butterfly species.
Ecology and Evolution, 3, 5167-5176.
Kopelman, N. M., Mayzel, J., Jakobsson, M., Rosenberg, N. A., and
Mayrose, I. (2015) Clumpak: a program for identifying clustering modes
and packaging population structure inferences across K. Molecular
Ecology Resources, 15, 179-1191.
Kozak, K. H., & Wiens, J. (2006) Does niche conservatism promote
speciation? A case study in North American salamanders. Evolution,
60(12), 2604-2621.
Kozak, K. M., McMillan, W. O., Joron, M., and Jiggins, C. D. 2018.
Genome-wide admixture is common across the Heliconius radiation.
bioRxiv, 414201, doi: https://doi.org/10.1101/414201
Leaché, A. D. and Bouckaert, R. R. (2018) Species trees and species
delimitation with SNAPP: a tutorial and worked example. Workshop on
population and speciation genomics, Český Krumlov
Leaché, A. D., Fujita, M. K., Minin, V. N., and Bouchaert, R. R. (2014)
Species delimitation using genome-wide SNP data. Systematic Biology, 63,
534-542.
Leigh, J. W., and Bryant, D. (2015) PopART: Full-feature software for
haplotype network construction. Methods in Ecology and Evolution, 6,
1110-1116.
Loera, I., Sosa, V., & Ickert-Bond, S. M. (2012). Diversification in
North American arid lands: Niche conservatism, divergence and expansion
of habitat explain speciation in the genus Ephedra. Molecular
phylogenetics and evolution, 65(2), 437-450.
MacDonald, Z. G., Nielsen, S. E., & Acorn, J. H. (2017) Negative
relationships between species richness and evenness render common
diversity indices inadequate for assessing long-term trends in butterfly
diversity. Biodiversity and conservation, 26(3), 617-629.
MacDonald Z.G., Dupuis J.R., Davis C.S., Acorn, J.H., Nielsen S.E.,
Sperling F.A.H. (2020). Gene flow and climate-associated genetic
variation in a vagile habitat specialist. Molecular Ecology.
Mace, G. M. (2004) The role of taxonomy in species conservation.
Philosophical Transactions of the Royal Society of London. Series B:
Biological Sciences, 359(1444), 711-719.
Magurran, A. E. (2013). Measuring biological diversity. John Wiley &
Sons.
Manel, S., Schwartz, M. K., Luikart, G., & Taberlet, P. (2003)
Landscape genetics: combining landscape ecology and population genetics.
Trends in Ecology & Evolution, 18(4), 189-197.
Mayden, R. L. (1997) A hierarchy of species concepts: The denouement in
the saga of the species problem. In Species: The units of biodiversity
(pp. 381–424). Chapman & Hall.
Mayr, E. (1942) Systematics and the origin of species. Columbia
University Press, New York.
Mayr, E. (1957) The species problem. American Association for the
Advancement of Sciences, Washington, DC.
Mayr, E. (1963) Animal Species and Evolution. Harvard University Press,
Cambridge, MA.
McHugh, A. Bierzychudek, P., Greever, C., Marzulla, T., Van Buskirk, R.,
and Binford, G., (2013) A molecular phylogenetic analysis of Speyeria
and its implications for the management of the threatened S. zerene
hippolyta. Journal of Insect Conservation, 17, 1237-1253.
McRae, B. H. (2006). Isolation by resistance. Evolution, 60, 1551-1561.
McRae, B. H., Beier P. (2007). Circuit theory predicts gene flow in
plant and animal populations. Proceedings of the National Academy of
Sciences, USA, 104, 19885-19890.
Miller, L. D., and Brown, F. M. (1981) Memoir No. 2: A catalogue/
checklist of the butterflies of America north of Mexico. Pp. 143. The
Lepidopterists’ Society.
Moeck, A. H. (1975) Geographic variability in Speyeria: comments,
records, and description of a new subspecies. Los Angeles: Entomological
Reprint Specialists.
Newton, L. G., Starrett, J., Hendrixson, B. E., Derkarabetian, S., &
Bond, J. E. (2020) Integrative species delimitation reveals cryptic
diversity in the southern Appalachian Antrodiaetus unicolor (Araneae:
Antrodiaetidae) species complex. Molecular Ecology.
Nguyen, L. -T., Schmidt, H. A., von Haeseler, A., and Minh, B. Q. (2015)
IQ-TREE: a fast and effective stochastic algorithm for estimating
maximum likelihood phylogenies. Molecular Biology and Evolution,
32:268-274.
Nosil, P. (2012). Ecological speciation. Oxford University Press.
Opler, P. A. and Warren, A. D. (2005) Lepidoptera of North America. 4.
Scientific names list for butterfly species of North America, North of
Mexico. Contributions of the C. P. Gillette Museum of Arthropod
Diversity, Colorado State University, Fort Collins, Colorado, USA.
Paris, J. P., Stevens, J. R., and Catchen, J. M. 2017. Lost in parameter
space: a road map for STACKS. Methods in Ecology and Evolution ,8 , 1360-1373.
Pebesma EJ, Bivand RS (2005). “Classes and methods for spatial data in
R.” R News, 5(2), 9–13.
Peterson, A. T. (2001). Predicting species’ geographic distributions
based on ecological niche modeling. The Condor, 103(3), 599-605.
Peterson, B. K., Weber, J. N., Kay, E. H., Fisher, H. S., and Hoekstra,
H. E. (2012) Double digest RADseq: an inexpensive method for de
novo SNP discovery and genotyping in model and non-model species.PLoS One , 7 , e37135.
Pelham, J. P. (2019) A catalogue of the butterflies of the United States
and Canada; with a complete bibliography of the descriptive and
systematic literature. The Lepidoptera Research Foundation/Journal of
Research on the Lepidoptera, Volume 40, Beverly Hills, CA, USA.
Phillips, S. J., Anderson, R. P., & Schapire, R. E. (2006). Maximum
entropy modeling of species geographic distributions. Ecological
modelling, 190(3-4), 231-259.
Pickrell, J. K., and Pritchard, J. K. (2012) Inference of population
splits and mixtures from genome-wide allele frequency data. PLoS
Genetics, 8, e1002967-17.
Poland, J. A., Brown, P. J., Sorrells, M. E., and Jannink, J. (2012)
Development of high-density genetic maps for barley and wheat using a
novel two-enzyme genotyping-by-sequencing approach. PLoS One ,7 , e32253.
Pritchard, J. K., Stephens, M., and Donnelly, P. (2000) Inference of
population structure using multilocus genotype data. Genetics, 155,
945-959.
R Core Team. (2017) R: A language and environment for statistical
computing. R Foundation for Statistical Computing, Vienna, Austria. URL
https://www.R-project.org/.
Rambaut, A. Drummond, A. J., Xie, D., Baele, G., and Suchard, M. A.
(2018) Posterior summarization in Bayesian phylogenetics using Tracer
1.7. Systematic Biology, 67, 901-904.
Reich, D. Thangaraj, K., Patterson, N., Price, A. L., and Singh, L.
(2009) Reconstructing Indian population history. Nature, 461, 489-494.
Revell, L. J. (2012) phytools: An R package for phylogenetic comparative
biology (and other things). Methods in Ecology and Evolution, 3,
217-223.
Riva, F., Campbell, E. O., Carroll, F., and Acorn, J. H. 2019.
Identification “by eye”: integrative character assessment informs
regional field identification of greater fritillary butterflies
(Nymphalidae: Speyeria). Journal of Insect Conservation 24, 259-267.
Rochette, N. C., Rivera-Colón, A. G., and Catchen, J. M. (2019) Stacks
2: analytical methods for paired-end sequencing improve RADseq-based
population genomics. Molecular Ecology, DOI:10.1111/MEC.15253
Sánchez-Ramírez, S., Rico, Y., Berry, K. H., Edwards, T., Karl, A. E.,
Henen, B. T., & Murphy, R. W. (2018). Landscape limits gene flow and
drives population structure in Agassiz’s desert tortoise (Gopherus
agassizii). Scientific reports, 8(1), 11231.
Schluter, D. (2001) Ecology and the origin of species. Trends in ecology
& evolution, 16(7), 372-380.
Schluter, D. (2009) Evidence for ecological speciation and its
alternative. Science, 323(5915), 737-741.
Scott, J. A., Kondla, N. G., and Spomer, S. M. (1998) Speyeria hesperis
and Speyeria atlantis are distinct species. Papilio New Series, 8, 1-32.
Sims, S. R. (2017) Speyeria (Lepidoptera: Nymphalidae) conservation.
Insects, 8, 45.
Sites, J. W., Jr., and Marshall, J. C. (2003) Delimiting species: a
Renaissance issue in systematic biology. Trends in Ecology and
Evolution, 18, 462-470.
Sonsthagen, S. A., Wilson, R. E., and underwood, J. G. (2017) Genetic
implications of bottleneck effects of differing severities of genetic
diversity in naturally recovering populations: An example from Hawaiian
coot and Hawaiian gallinule. Ecology and Evolution, 7, 9925-9934.
Sperling, F. A. H. (2003) Butterfly molecular systematics: from species
definitions to higher level phylogenies. Pp. 431-458. IN Boggs, C.,
Erlich, P. and Watt, W. (Eds). Ecology and Evolution Taking Flight:
Butterflies as model study systems. University of Chicago Press,
Chicago.
Stanton, D. W. G., Frandsen, P., Waples, R. K., Russo, I-R. M.,
Orozco-terWengel, P. A., Tingskov Pedersen, C-E., Siegismund H. R., and
Bruford, M. W. (2019) More grist for the mill? Species delimitation in
the genomic era and its implications for conservation. Conservation
Genetics, 20, 101-113.
Storfer, A., Murphy, M. A., Spear, S. F., Holderegger, R., & Waits, L.
P. (2010). Landscape genetics: where are we now?. Molecular Ecology,
19(17), 3496-3514.
Thompson, E., Baumsteiger, J., and Hill, R. I. (2019) Phylogenomic
analyses clarify true species within the butterfly genus Speyeria
despite evidence of a recent adaptive radiation. Insects, 10, 209-224.
Thorpe, R. S., Surget‐Groba, Y., & Johansson, H. (2008) The relative
importance of ecology and geographic isolation for speciation in anoles.
Philosophical Transactions of the Royal Society B, Biol. Sci., 363,
3071-3081.
Thorpe, R. S., Surget-Groba, Y., & Johansson, H. (2010) Genetic tests
for ecological and allopatric speciation in anoles on an island
archipelago. PLoS Genetics, 6(4).
van Etten J (2017) “R Package gdistance: Distances and Routes on
Geographical Grids.” Journal of Statistical Software, 76(13), 21.
Van Valen, L. (1976) Ecological species, multispecies, and oaks. Taxon,
233-239.
Vargas, O. M., Ortiz, E. M., Simpson, B. B. (2017) Conflicting
phylogenomic signals reveal a pattern of reticulate evolution in a
recent high-Andean diversification (Asteraceae: Astereae:
Diplostephium). New Phytologist, 214, 1736-1750.
Wagner, C. E., Keller, I., Wittwer, S., Selz, O. M., Mwaiko, S.,
Greuter, L., Sivasundar, A., and Seehausen, O. (2013) Genome-wide RAD
sequence data provide unprecedented resolution of species boundaries and
relationships in the Lake Victoria cichlid adaptive radiation. Molecular
Ecology, 22, 787-798.
Wang, Y. H., Yang, K. C., Bridgman, C. L., & Lin, L. K. (2008) Habitat
suitability modelling to correlate gene flow with landscape
connectivity. Landscape Ecology, 23(8), 989-1000.
Wang, I. J., Glor, R. E., & Losos, J. B. (2012) Quantifying the roles
of ecology and geography in spatial genetic divergence. Ecology letters,
16(2), 175-182.
Warren, A. D., Davis, K. J., Grishin, N. V., Pelham, J. P., and
Stangeland, E. M. (2012) Interactive listing of American butterflies
<www.butterfliesofamerica.com>
Werren, J. H., Baldo, L., and Clark, M. E. (2008) Wolbachia: master
manipulators of invertebrate biology. Nature Reviews Microbiology, 6,
741-751.
Wiens, J. J. (2004) What is speciation and how should we study it? Am.
Nat. 163, 914–923.
Wilkins, J. S. (2009) Species: A history of the idea. Berkeley (CA):
University of California Press.
Zimmermann, N. E., Edwards Jr, T. C., Graham, C. H., Pearman, P. B., &
Svenning, J. C. (2010) New trends in species distribution modelling.
Ecography, 33(6), 985-989.