List of symbols A area = area-based photosynthetic rate, Amass = mass-based photosynthetic rate, gsw area = area-based stomatal conductance, gsw mass = mass- based stomatal conductance, E area = area-base transpiration rate, E mass = mass-based transpiration rate, SLA= specific leaf area, Nconcentration = nitrogen concentration (g/kg), Pconcentration = phosphorus concentration (g/kg), ΨPD = pre-dawn water potential, ΨMD = mid-day water potential, WUE = water use efficiency, WUEi = Intrinsic water use efficiency, PNUE =photosynthetic nitrogen use efficiency, PPUE = photosynthetic phosphorus use efficiency, N area = area-based leaf nitrogen concentration (g m-2), P area = area-based leaf phosphorus concentration (g m-2), Chl area = area-based leaf total chlorophyll concentration (mg g-1).