Acknowledgements
We would like to thank the landowners in Ambodiala and Antsikory who let us survey their lands. We also thank our field assistants Romual Randrianantenaina and Erosiniot Befidimanana for their great support during fieldwork and James Herrera for his feedback that improved the quality of the first draft of this manuscript. We also thank Miguel Vences from Evolutionary Biology laboratory for genetic identification of the frog species. This study was financially supported by the Volkswagen Stiftung within the project ‘Diversity Turn in Land Use Science’ (Grant number 11-76251-99-35/13 (ZN3119)) which is a collaboration between the University of Gottingen, the University of Antananarivo, Madagascar National Parks and the University of CURSA (University Center of SAVA Region). TRF was supported by the German Academic Exchange Service (DAAD) within the ‘Partnerships for Supporting Biodiversity in Developing Countries’ initiative (Project Nr.57449386). We collected data under the research permit N°254/18/MEEF/SG/DGF/DSAP/SCB.Re granted by the Ministry for Water, Ecology and Forest (MEEF), Antananarivo.