ACKNOWLEDGMENTS
This work was supported by the grants from National Natural Science Foundation of China (31870296, 31700277), Natural Science Foundation of Henan (No. 182300410063), Nanhu Scholars Program for Young Scholars of Xinyang Normal University, the Shanghai Science and Technology Committee (18DZ2260500, 17DZ2252700), and the Collaborative Innovation Center for Modern Crop Production to H.Z.