ACKNOWLEDGEMENTS
We are grateful to L. Wu for assistance in the field. We thank Proof-Reading-Service.com Ltd in Letchworth Garden city for the helpful feedback on a previous draft of the manuscript. This study supported by Strategic Priority Research Program of the Chinese Academy of Sciences (XDB31030000), the National Key R&D Program of China (grand No. 2017YFC0505802), National Natural Science Foundation of China (No. 31370437), National Natural Science Foundation of China (No. 41371078) and Chinese Forest Biodiversity Monitoring Network.
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