Acknowledgment
This work was financially supported by Natural Science Foundation of China (Grants 31972378), National Key Research and Development Program of China (2019YFD1000100), Agricultural Variety Improvement Project of Shandong Province (2019LZGC007), and Ministry of Agriculture of China (CARS‐27). We would like to thank Professor Fang-Fang Ma of Shandong Agricultural University for generous help in organic acids determination experiments and analysis of the data.
Author Contributions
Y.-J. H., X.-F. W., W.-J. S. conceived the study and wrote the manuscript. W.-J. S., X.-L. J., J.-C. Z., Z.-Q. F. performed mass spec, W.-J. S, J.-C. Z., Z.-Q. F. performed metal content and nitrogen content analysis experiments, W.-J. S., Z.-Q. F., C.-X. Y., W.-J. H. performed phenotypic analysis and field experiments, W.-J. S., X. W., Z.-Q. F. performed the qPCR and the RNA-Seq experiments, W.-J. S., X.-L. J., Z.-Q. F., C.-X. Y. performed the chlorophyll, pH and FCR analysis experiments.