Acknowledgment
This work was financially supported by Natural Science Foundation of
China (Grants 31972378), National Key Research and Development Program
of China (2019YFD1000100), Agricultural Variety Improvement Project of
Shandong Province (2019LZGC007), and Ministry of Agriculture of China
(CARS‐27). We would like to thank Professor Fang-Fang Ma of Shandong
Agricultural University for generous help in organic acids determination
experiments and analysis of the data.
Author Contributions
Y.-J. H., X.-F. W., W.-J. S. conceived the study and wrote the
manuscript. W.-J. S., X.-L. J., J.-C. Z., Z.-Q. F. performed mass spec,
W.-J. S, J.-C. Z., Z.-Q. F. performed metal content and nitrogen content
analysis experiments, W.-J. S., Z.-Q.
F., C.-X. Y., W.-J. H. performed phenotypic analysis and field
experiments, W.-J. S., X. W., Z.-Q. F. performed the qPCR and the
RNA-Seq experiments, W.-J. S., X.-L. J., Z.-Q. F., C.-X. Y. performed
the chlorophyll, pH and FCR analysis experiments.