Acknowledgements
This research was funded by the National Key Research and Development Program (2016YFD0100201, 2018YFD0201006), the National Natural Science Foundation of China (Grant Nos. 31871646 and 31771821), the MOE Program for Changjiang Scholars and Innovative Research Team in University (PCSIRT_17R55), the Fundamental Research Funds for the Central Universities (KYT201801), and the Jiangsu Collaborative Innovation Center for Modern Crop Production (JCIC-MCP) Program.