Environmental stress level Heterogeneous area Heterogeneous area Heterogeneous area Transitional area Transitional area Transitional area Transitional area Transitional area Homogeneous area Homogeneous area Homogeneous area
Emin Emax Richness Emin Emax Richness Richness Emin Emin Emax Richness
All optima niche 0 6 108.4±4.7b 0 4 4 117.7±3.0a 117.7±3.0a 0 2 95.2±5.2c
Half optimal niche 1 7 91.3±4.3b 1 5 5 97.0±4.1a 97.0±4.1a 1 3 87.8±2.3b
No optimal niche 3 9 51.4±3.0a 3 7 7 48.3±3.5ab 48.3±3.5ab 3 5 44.6±4.9b
Highest stress 4 10 34.4±3.8a 6 10 10 19.5±2.5b 19.5±2.5b 8 10 9.5±2.2c