Both tight junctions and adherens junctions are connected to the actomyosin cytoskeleton, and tightly regulated actomyosin dynamics are critical not only for junction formation, but also for structure and function of tight junctions and adherens junctions during epithelial tissue homeostasis, and morphogenesis \cite{Lecuit2015,Rodgers2011,Takeichi2014}. Similar to the contractile ring that forms during cell division Rho GTPases are critical for proper dynamic organization and contractility of junctional actomyosin. When misregulated, abnormal junctional F-actin polymerization or Myosin II contraction can drive pathological conditions \cite{Ivanov_2010,Marchiando_2010}.