Abstract
The Equilibrium Theory of Island Biogeography (ETIB) is a widely applied
dynamic theory proposed to explain why islands have coherent differences
in species richness. The development of the ETIB was temporarily
challenged by the alternative static Theory of Ecological Impoverishment
(TEI), which suggests that the number of species on an island is
determined by its number of niches or habitats. With no clear evidence
relating species richness to the number of niches, the TEI was almost
abandoned in favour of the ETIB. Here, we show that the number of
climatic niches on islands is an important predictor of the species
richness of plants, herpetofauna and land birds, and we propose a new
model called the General Theory of Island Biogeography (GTIB) that
merges the ETIB and TEI theories. When we test our GTIB for resident
land birds in Krakatau Islands it reveals a good correspondence with
observed species richness, immigration and extinction rates.