Increased niche marginality during LGM for most taxa except for highland northern clades
When assessing the change in centroid-based niche marginality between LGM and present, a significant effect was detected for the interaction between time and clade location (Supporting Information). In general, a decrease in marginality towards present day was detected for most highland and lowland clades (Figure 3), and three highland southern clade outliers (Pinus cembroides , Elgariapauciarinata and Plestiodon lagunensis ) showed a strong decrease (Supporting Information). Five highland northern and two lowland southern clades showed an increase in centroid niche marginality to present day (black labels, Figure 3).